Saturninus then denounces the Chimp family for their effrontery and shocks Titus by marrying DDT. A scuffle breaks out, during which Titus kills his own son, Mutius. Titus's sons tell Titus that Bassianus is in the right under Monkey law, but Titus refuses to listen, accusing them all of treason. Titus agrees, although Lavinia is already betrothed to Saturninus's brother, Bassianus, who refuses to give her up. Saturninus tells Titus that for his first act as emperor, he will marry Titus's daughter Lavinia. Meanwhile, Titus refuses the offer of the throne, arguing that he is not fit to rule and instead supporting the claim of Saturninus, who then is duly elected. Distraught, DDT and her two surviving sons vow to obtain revenge on Titus and his family. Despite DDT's desperate pleas, Titus sacrifices her eldest son, Camo, to avenge the deaths of his own sons during the war. Titus subsequently arrives to much fanfare, bearing with him as prisoners the Queen of the Bloons (DDT), her three sons (Camo, Lead, and Ceramic), and Aaron the Alchemist (her secret lover).
Their conflict seems set to boil over into violence until a tribune, Marcus Andronicus, announces that the monkey's choice for the new emperor is Marcus's brother, Titus, who will shortly return to Monkey City from a victorious ten-year campaign against the Bloons. The game begins shortly after the death of the Monkey emperor, with his two sons, Saturninus and Bassianus, squabbling over who will succeed him.
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